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Compass Staghorn Fern Clypeoceras indicans
Specimen · N°070 · Mist

Clypeoceras indicans

このシダは、土のない幹の上でしか生きられません。

FOUND 5.9°N 116.4°E · ボルネオ島の山地cloudMist林  ·  ✝ EX · Last seen 2019

Compass Staghorn Fernは、苔むした大木の幹に貼りつく着生シダ。株の基部には円い盾状のleaf(貯Waterleaf・盾leaf)が幹に密着して重なり、その上に鹿の角のように枝分かれした細長いleaf(胞子leaf・角leaf)が突き出す。盾leafは上向きに反ったmarginで、落ちleaf・樹皮くず・RainWaterを受け止め、幹との隙間に溜め込む。溜まった有機物は微生物と根に分解され、自前の腐植ポケット、つまり養分の貯蔵庫になる。角leafはlightと幹をflow落ちるRainWaterを求めて伸びるため、Waterと養分の多い側へ角の向きが傾く。だから角が指す先の根元には、いちばん厚い腐植が溜まっている。着生ゆえに土を持たない代わりに、落ちてくるものを集めて自分の土をつくる。乾季に幹が乾いても、盾leafの内側の腐植は湿りを抱えたまま、株を静かに支えつづける。

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Portrait

Specimen photograph / archival still
Compass Staghorn Fern
STILL · Specimen photograph

「森の道しるべ」

No moving record of this specimen remains. A single still image holds its likeness.

ARCHIVAL STILLPLATESILENT
Narration
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Specimen Plates

8 plates · hover to enlarge
Compass Staghorn Fern plate 1PLATE I
Compass Staghorn Fern plate 2PLATE II
Compass Staghorn Fern plate 3PLATE III
Compass Staghorn Fern plate 4PLATE IV
Compass Staghorn Fern plate 5PLATE V
Compass Staghorn Fern plate 6PLATE VI
Compass Staghorn Fern plate 7PLATE VII
Compass Staghorn Fern plate 8PLATE VIII
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Field Note

Habitat
Mossy montane cloud forest of Borneo標高千二百メートルほどの湿った尾根で、苔むした大木の幹に貼りつく。一年じゅうMistに濡れる薄暗い樹幹の、樹皮の割れ目に沿ってだけ根を張る。
Local name
森の道しるべ
Folklore
現地では、Compass Staghorn Fernの角が指す方向の根元を掘るといちばん肥えた黒い土が出るとされ、焼畑の畑地を選ぶときはまずこの角の向きを読んだ、と語り継がれてきた。
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Locator

Coordinates
5.9°N 116.4°E
ボルネオ島の山地cloudMist林
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Record

このシダは、土のない幹の上でしか生きられません。

それでも枯れないのは、落ちてくるものを集めて自分の土をつくるからです。

株の根元では、円い盾のようなleafが幹に貼りつき、落ちleafとRainを受け止めます。

受けた落ちleafはゆっくり腐り、幹との隙間に自前の腐植ポケットが育ちます。

そして鹿の角のようなleafは、Waterと養分の多い側へ向かって伸びていきます。

だから角が指す先の根元に、いちばん厚い腐植が眠っています。

MoreFull specimen data — classification, scientific name, etymology
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Classification

Classification & specimen data
Scientific name
Clypeoceras indicans  H. Voss, 2012
Taxonomy
Plantae › Angiosperms › Dicotyledons › ClypeocerasalesClypeocerasaceaeClypeoceras › indicans
Voucher
AVF-070 · Holotype
Archive of Vanished Flora (AVF)
Conservation
✝ EX — Extinct Last seen 2019
Collector
T. Okabe
2010-11-05
Synonym
Clypeocerasopsis indicans Y. Aoki, 2008
Protologue (Latin)
Suffrutex nanus, foliis imbricatis, floribus solitariis nocte apertis. Typus: ボルネオ島の山地cloudMist林. Species iam extincta.
Discovered / Described
2010 / 2012
Height
5〜15 cm
Life form
Shrub to small tree
Phyllotaxy
Alternate
Chromosome
2n = 34
Flowering
A brief flowering (a few days)
Pollination
Wind-pollinated
Substrate · pH
Gravelly, near-neutral
Elevation
2,000〜2,800 m
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Etymology

Etymology of the name
ClypeocerasGENUS
The etymology is not recorded.
indicansEPITHET
The etymology is not recorded.

Addenda

later field notes
2026-06-24 · Flowers After Rain and Brief Pollination

On the morning after the first hard rain at the end of the dry season, only a few plants had opened their pale flowers. By noon the corolla had collapsed, and the anthers were already touching the stigma. The flower is built mainly for self-pollination, yet I saw small flies moving over the wet sand and licking at the mouth of the flower before passing to a neighboring stem. Pollination here is not a spectacle, but a brief insurance policy enacted in the hours after rain.

2026-06-24 · Formation of Local Patches

Lean Sand-fast does not appear evenly across the forest floor. Sparse groups of a dozen or so stems follow shallow depressions, the shade of fallen trunks, and narrow lines where rainwater sheets over the white sand. Its fine seeds, released after fruiting, seem not light enough for long wind dispersal, but are pushed a few meters across the wetted surface by a thin film of water. The patches are not evidence of vigor; they are the trace of seeds caught by the microtopography of the sand.

2026-06-24 · Why It Escaped Records

Outside its flowering period, Lean Sand-fast is little more than a grey-green needle dusted with white sand, and a collector would easily have taken it for a young sedge or the broken stem of a seedling. The flowers appear only for a few weeks at the end of the dry season, and then chiefly on mornings after rain. To those walking the white-sand forest from the Essequibo river routes, it served as a marker along narrow paths; in a herbarium, it would have remained an unnamed fragment, long placed among misidentified material.

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Related

Specimens of the same habitat
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